For a full list of works by Dr. Robert Drewes, please contact the Herpetology department.


  • A Field Guide to the Reptiles of East Africa (2002)
  • Pocket Guide to the Reptiles and Amphibians of East Africa (2006)
  • Environmental Physiology of the Amphibians (2008)

Scientific Articles and Publications

  • Drewes, R. C. , M. S. Hedrick, S. S. Hillman and P. C. Withers Unique role of skeletal muscle in vertical lymph movement in anurans In press. Journal of Experimental Biology
  • Uyeda, J., R. C. Drewes and B. M. Zimkus. The California Academy of Sciences Gulf of Guinea Expeditions (2001, 2006) VI. A new species of Phyrynobatrachus from the Gulf of Guinea Islands and a reanalysis of Phrynobatrachus dispar and P. feae (Anura: Phrynobatrachidae). 2007, Proceedings of the California Academy of Sciences 58:367-385.
  • Measey, J. G., M. Vences, R. C. Drewes, Y. Chiari, M. Melo and B. Bourles. Freshwater paths across the ocean: Molecular phylogeny of Ptychadena newtoni gives insights into amphibian colonization of oceanic islands. 2007. Journal of Biogeography 34:7-20.
  • Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sa, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green and W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297:1-3
  • Hillman, S. S., P. C. Withers, M. S. Hedrick and R. C. Drewes 2005. Functional roles for the compartmentalization of the subcutaneous lymphatic sacs of anuran amphibians. Physiological and Biochemical Zoology 78: 515-523.
  • Drewes, R. C. & R. E. Stoelting 2004. The California Academy of Sciences Gulf of Guinea Expedition (2001) II. Additions and corrections to our knowledge of the endemic amphibians of São Tomé and Príncipe. Proceedings of the California Academy of Sciences. 55 (31):573-587.
  • Drewes, R. C. & J. A. Wilkinson 2004. The California Academy of Sciences Gulf of Guinea Expedition (2001) I. The taxonomic status of the genus Nesionixalus Perret (Anura: Hyperoliidae), treefrogs of São Tomé and Príncipe, with comments on the genus Hyperolius. Proc. Calif. Acad. Sci. 55 (20):395-407.
  • Wilkinson, J. A., R. C. Drewes & O. L. Tatum 2002. A molecular phylogenetic analysis of the family Rhacophoridae with an emphasis on the Asian and African genera. Molecular and Phylogenetic Evolution 24:265-273
  • Wieczorek, A. M., R. C. Drewes & A. Channing 2001. Phylogenetic relationships within the Hyperolius viridiflavus complex (Anura, Hyperoliidae), and comments on taxonomy. Amphibia-Reptilia 22:155-166.
  • Wieczorek, A. M., R. C. Drewes & A. Channing 2000. Biogeography and evolutionary history of Hyperolius species: application of molecular phylogeny. Journal of Biogeography 27:1231-1243
  • Drewes, R. C. & J.- L. Perret 2000. A new species of giant, montane Phrynobatrachus (Anura: Ranidae) from the central mountains of Kenya. Proceedings of the California Academy of Sciences 52:55-64.
  • Emerson, S. B., C. Richards, R. C. Drewes & K. M. Kjer 2000. On the relationships among ranoid frogs: a review of the evidence. Herpetologica 56:209-23.
  • Hillman, S. S., P. C. Withers & R. C. Drewes 2000. Correlation of ventricle mass and dehydration tolerance in amphibians. Herpetologica 56(4): 413-420.
  • Drewes, R. C. A new species of treefrog from the Serengeti National Park, Tanzania (Anura: Hyperoliidae: Hyperolius). 1997. Proceedings of the California Academy of Sciences 49: 439-446.
  • Drewes, R. C. & R. Altig 1996. Anuran egg predation and heterocannibalism in a breeding community of East African frogs. Tropical Zoolog. 9:333-347.
  • Drewes, R. C. & J. V. Vindum. 1994. Amphibians of the Impenetrable Forest, southwest Uganda. Journal of African Zoology 108:55-70.
  • Drewes, R. C. 1985. A case of paraphyly in the genus Kassina Girard, 1853. South African Journal of Science. 81 186- 191.
  • Drewes, R.C. 1984. A phylogenetic analysis of the Hyperoliidae (Anura): treefrogs of Africa, Madagascar and the Seychelles Islands. Proceedings of the California Academy of Sciences. 139: 1-70.
  • Drewes, R.C., S. S. Hillman, R. W. Putnam & O. Sokol. 1977. Water, nitrogen and ion balance in the desert treefrog Chiromantis petersi Boulenger (Anura: Rhacophoridae) with comments on the structure of the integument. Journal of Comparative Physiology 116(39): 257-267.

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Dr. Drewes on IBSS

Learn more about Dr. Drewes' research, publications, and current projects by visiting the research side of our website, home to the Academy's Institute for Science and Sustainability.

Herpetology at the Academy

The Academy's Herpetology collection of amphibians and reptiles is one of the 10 largest in the world, containing more than 309,000 cataloged specimens from 175 countries. Learn more about the department's staff, research, and expeditions.