Curator of Entomology, Emeritus
Systematics, biogeography, evolution, and natural history of carabid beetles (Coleoptera: Carabidae).
Changes in the altitudinal distributions of montane carabid beetles as indicators of climate change.
Biogeography, ecology, and evolution of high-altitude, montane organisms and faunas.
General aspects of biogeography and evolution.
General principles and methods of systematics.